Latent life- dormancy:
Importance and types of dormancy,
Dormancy
As stated
earlier, seeds of some plants, fail to germinate even if placed under all the
favorable conditions such as moisture, air, suitable temperature and proper
sequence of light and dark. In most of
the cases, the germination may be delayed for days, weeks, months or even
years. Such seeds are said to be in the state of dormancy. Dormancy may be
defined as any rest period or reversible interruption of the phenotypic
development of an organism or a state, in which seeds fail to germinate under
conditions of moisture, temperature, and oxygen favorable for vegetative growth
Importance
The phenomenon
of dormancy is widely spread in living organisms. Bacteria, fungi, insects, and
animals also undergo periods of reduced growth activity i.e. dormancy. Looking
to the differences in growth habits, nutrition, temperature and water
regulation mechanism, and metabolism of these groups, it is unlikely that the
same basic mechanism is involved in establishing the dormant condition in these
different groups.
The significance of dormancy may be considered as-
·
It
enables them to endure periods of unfavorable environmental conditions.
·
In
temperate zones, it helps the seeds to pass through the severe colds.
·
In
tropical regions, the impermeable seed coats ensure good chances of survival.
·
Dormant
seeds and organs in perennial plants resist unfavorable conditions for their
development
·
It
enables the seed for long distance dispersals.
·
The
seeds form a measure of the quantity and duration of rainfall, both of which
determine the amount of soil- moisture available for plant growth.
·
The
dormancy of seeds of cereals is most important to mankind. If these seeds
germinate immediately after harvest, they will be quite useless to mankind.
Types
There are
several classifications of seed dormancy, proposed by different authors.
The main types of dormancy are:
The Imposed Dormancy
(Quiescence)
Germination
needs favorable environmetal conditions like sufficient water, proper
temperature, etc. in absence of such conditions seed does not germinate or
becomes dormant which is sometimes considered as inhibition. This type of
inhibition is known as imposed dormancy (Quiescence).however, under favorable
conditions this kind of dormancy disappears and the seed germinates
immediately.
The Innate
(Deep) Dormancy
Sometimes a
seed unable to germinate due to structural, chemical or physiological
properties of the seed itself, this phenomenon is known as Innate (deep)
dormancy and is distinguished as physical, chemical, morphological or physiological
types of dormancy. These terms describe the main mechanisms that are
responsible for the inability of embryo to grow: physical barrier preventing
water and/or oxygen access, presence of growth inhibitors, and morphological or
physiological immaturity of embryo. These mechanisms can operate separately or
in various combinations.
The fully
dormant condition of a seed is a gradual process and is not attained suddenly.
During the entire process, there may be following three phases of dormancy.
(I)Predormancy or early rest :
During this phase, the dormant organ has
capacity to resume growth by various treatments i. e. , capacity of germination
or growth is not completely lost.
(II) Full dormancy or mid rest :
When a seed or
organ becomes completely dormant and germination or growth can not be induced
immediately by changing environmental conditions.
(III) Post dormancy or after rest :
When a dormant seed or organ gradually emerges from full dormancy and in it the
germination or growth can be induced by
changing environmental conditions.
Complex types of dormancy
These are
caused by several different mechanisms, which make germination difficult or
hinder it altogether. These mechanisms act as a set of barriers, which prevent
germination.
The dormancy
may be true, relative or secondary.
(I) True dormancy :
when in a seed
or organ , the germination or growth can not be induced under any set of
environmental conditions.
(II) Relative dormancy :
when in a seed,
the germination can be induced under specific conditions even at the time of
its deepest dormancy.
(III) Secondary dormancy :
when a seed has
not fully emerged from dormancy and is again thrown back into full dormancy by
certain environmental conditions.
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